CLEMSON ­— So many times during the last two seasons, little mistakes resulted in big problems for the Clemson defense. Little things, like a linebacker playing on a blocker’s inside shoulder rather than staying outside, thus losing leverage and leading to a big play.

But on second-and-11 play in the second quarter at Wake Forest, Clemson linebacker Jonathan “Tig” Willard stayed on the outside shoulder of a Wake blocker. It was tempting to move inside because that’s where Wake’s Lovell Jackson appeared to be headed. But Willard stayed disciplined, he stayed outside. So when Jackson was funneled outside, Willard was there to make an open-field tackle.

“I couldn’t let him get outside so I had to fight to stay outside the whole time,” Willard said. “Last year, I didn’t know leverage-wise where I was supposed to be.”

So many times over the last two seasons, Clemson defenders had failed to make impact plays. Clemson linebackers recorded just 15 tackles for loss last season.

But on the following third down play at Wake, Willard blitzed. The senior dipped his shoulder below the hands of a Wake offensive lineman and accelerated into quarterback Tanner Price, leveling him for a sack.

After so much defensive inconsistency, Willard has become a consistent performer, leading what Clemson hopes is a defensive turnaround with a team-best 62.

“It’s football 101, understanding not just what I’m supposed to do, but why,” Clemson defensive coordinator Brent Venables said. “It’s muscle memory.”

Willard’s stated goal is a 100-tackle season. He’s coming off back-to-back, double-digit tackle performances. He has three tackles for loss in the last two weeks and leads Clemson with 5˝ tackles for loss on the season.

Over the last four games, Clemson has improved from 111th in the country in yards allowed per rush (5.4) to 94th (4.8). Clemson’s total defense ranking improved from 95th overall to 85th this week. Leading the improvement is Willard.

“He’s a guy that’s played a lot of football. A young man that cares deeply, very selfless and he’s maximized his opportunities,” Venables said.

The 6-foot-2, 220-pounder former high school wrestling star has played with a torn ulnar collateral ligament in his elbow that affects the arm’s range of motion. In baseball, the injury would require Tommy John surgery.

“Wrestlers are tough,” Venables said. “I love wrestlers. They have something a lot of guys don’t have.”